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n.
the discipline dealing with the art or science of applying scientific knowledge to practical problems; "he had trouble deciding which branch of engineering to study"
n.
a perceptible indication of something not immediately apparent (as a visible clue that something has happened); "he showed signs of strain"; "they welcomed the signs of spring"
n.
an ability to understand the meaning or importance of something (or the knowledge acquired as a result); "how you can do that is beyond my comprehension"; "he was famous for his comprehension of American literature"